Central American montane forests

  • Category: Forest Biomes.
  • Geographic Location: Central America.
  • Summary: Home to many unique habitats, this cloud-forest biome represents an evolutionary crossroads of montane species from North and South America.

Montane forests occur in isolated patches in a mosaic along the mountaintop region from southern Mexico to northern Nicaragua. Predominantly cloud forests—marked by persistent seasonal or frequent low-level clouds—this biome is unique in representing part of a temperate dispersal corridor (together with Talamancan montane forests) between North and South America, where many elements from the north and south have mixed. The region has very high endemism (species found nowhere else on Earth) among plants, invertebrates, and vertebrates.

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The biome consists of at least 40 small habitat islands occurring in a chain along the Sierra Madre del Sur in Mexico, the Sierra de las Minas in Guatemala, and isolated regions of Honduras, El Salvador, and Nicaragua, most of it poorly represented by the national protected-areas systems of these countries. Montane elements typically occur at elevations above 4,921 feet (1,500 meters) and extend to more than 13,123 feet (4,000 meters) along the highest peaks. Above 9,843 feet (3,000 meters), forests become scrubby, and temperature extremes can be dramatic, resulting in unique alpine-like elements such as grasslands.

From 4,921 to 9,843 feet (1,500 to 3,000 meters), cloud forests are dominated by conifers and oaks, forming the pine-oak forest belt. Within these formations are bromeliad associations and bamboo stands. The origins of these mountains are diverse, ranging from Paleozoic sedimentary and metamorphic rocks to much more recent volcanic (some still active) formations dating back to the Pliocene Epoch.

Vegetation

The vegetation typical of this biome is dominated by components from the north with a mix of elements from the south that worked their way north following the closure of the Central American land bridge. The dominant natural cover type is pine-oak forests, and much of the land has been converted to human-dominated agricultural and urban matrices. This ecosystem represents the southernmost limit of native pine forests.

Among the unique features of the vegetation is high species endemism, reaching as high as 70 percent in some areas. The isolated mountain peaks along the central spine of northern Central America are not only isolated from one another, but also represent unique climatic gradients moving up from lowland rainforests to high, sometimes-snow-covered peaks. The pine-oak forests typically are a mix of conifers (Abies, Cupressus, Juniperus, and Taxus) and broadleaf evergreens such as oak (Quercus). This biome represents the southern limit of the conifer's native range, whereas oaks continue southward to the Talamancan montane forests and beyond.

The climate is temperate and precipitation is high, which is usual for tropical montane systems, and is mostly due here to the narrow extent of land that Mesoamerica represents. Weather patterns create unique microclimates, and the mountain systems give way to rain shadows and other effects that influence both flora and fauna. Precipitation typically is 79 to 157 inches (2,000 to 4,000 millimeters) per year, much of it occurring as rainfall but some as cloud drip. These cloud forests form an extremely high diversity of epiphytic species such as orchids, bromeliads, and many others that depend on—and can live only on—atmospheric moisture. Above 9,843 feet (3,000 meters), frost is typical, and plant communities become limited to frost-tolerant species.

Avian Life

Bird diversity and endemism are particularly high in this biome. Among the flagship species is the majestic horned guan (Oreophasis derbianus), which is increasingly threatened due to loss of its cloud-forest habitats. Another charismatic species is the resplendent quetzal (Pharomacrus mocinno), prized by bird-watchers and one featured in the art and culture of many local pre-Hispanic and current indigenous peoples.

Other species in the biome include the endangered ocellated quail (Cyrtonyx ocellatus); pink-headed warbler (Ergaticus versicolor); azure-rumped tanager (Tangara cabanisi); and the extinct Atitlan grebe (Podilymbus gigas), which disappeared in 1986. The biome has 50 Important Bird Areas, established to identify conservation priorities.

The area is also important to migrating birds. The area is characterized by local migrations and intercontinental migrations. More than 20 species from North America winter in this habitat. The area is also important to migrating butterflies.

Other Animals

Like birds, mammals are characterized by high diversity, but rates of endemism are lower than for birds or plants. One of the most interesting mammal species of the region is the very small volcano rabbit (Romerolagus diazi), native to Mexico and endangered. Also present are threatened species such as the Central American spider monkey (Ateles geoffroyi), the Mexican long-nosed bat (Leptonycteris nivalis), and the Baird's tapir (Tapirus bairdii). The biome represents one of the last habitats for these species across their range.

The area's amphibians and reptiles are unique, with diversity decreasing with elevation. One of the most remarkable characteristics is the high number of salamander and frog species. At least 28 threatened species occur across the biome, and the actual number probably is larger, considering the low documentation efforts in most of the countries. At least 13 species are critically endangered, including nine frogs and four salamanders. The decline of amphibians in this biome is associated with the threat from the chytrid fungus infection, which affects this group mostly at higher elevations. Climate change also seems to be an important issue for amphibian species in this biome, with altered temperature and moisture patterns leading to the necessity for some species to transit to new localities to find their comfort zones.

Effects of Human Activity

Due to their biological richness and their isolation, many of these mountain peaks have been declared protected areas, but little environmental enforcement is extended to them. Typically, however, the cloud forests are not only habitats. They have become isolated as much of the lowlands that previously spanned the mountains has been converted to agriculture and other human-dominated environments. Little connectivity now exists between fragments, so species have little dispersal ability. A secondary effect is that many species in this area, especially birds, rely on elevational migration routes that have become fragmented, if not completely altered.

The largest remaining forest block in the biome is Guatemala's Sierra de las Minas, which covers more than 4.9 million acres (2 million hectares). Although protected, it is also the only remaining native habitat of sufficient size to maintain all native species and ecological processes. Although many of the other montane areas within this biome are protected, most are remnants of their former distribution and today are isolated fragments of forest in a sea of agriculture. Typically, lower elevations are more highly converted, with intactness increasing with elevation—exactly the opposite trend required for biodiversity. Even though this biome is fairly intact in general, the areas with the highest biodiversity are the most at risk.

Bibliography

Baumbach, Lukas, et.al. "Climate Change May Induce Connectivity Loss And Mountain Extinction in Central American Forests." Communications Biology, vol. 4, no. 869, 15 Jul. 2021, doi.org/10.1038/s42003-021-02359-9. Accessed 1 Dec. 2024.

Halffter, G. “Biogeography of the Montane Entomofauna of Mexico and Central America.” Annual Review of Entomology 32, no. 1 (1987).

Land, H. C. Birds of Guatemala. Wynnewood, PA: Livingston, 1970.

Reid, F. Field Guide to the Mammals of Central America and Southeast Mexico. New York: Oxford University Press, 2009.

Rich, P. V. and T. H. Rich. “The Central American Dispersal Route: Biotic History and Paleogeography.” In D. H. Janzen, ed. Costa Rican Natural History. Chicago: University of Chicago Press, 1983.

Schipper, Jan. “Central American Montane Forests.” One Earth, www.oneearth.org/ecoregions/central-american-montane-forests/#:~:text=More%20than%2020%20bird%20species,resplendent%20quetzal%2C%20Pharomacrus%20mocinno%20mocinno. Accessed 1 Dec. 204.

Sutton, Susan Y. “Nicaragua.” In David G. Campbell and H. David Hammond, eds. Floristic Inventory of Tropical Countries. Bronx, NY: New York Botanical Garden, 1988.