Magellanic subpolar forests

  • Category: Forest Biomes.
  • Geographic Location: South America.
  • Summary: This subpolar realm, also known as Nothofagus forests, extends from Tierra del Fuego at the southern tip of South America into Chile and Argentina; it is a refuge for Antarctic flora.

The subpolar Nothofagus forests extend along the base of the southern Andes Mountains and across the Chilean archipelago, bounded on the south and west by the ocean; on the east by the dry Patagonian steppe; and on the north by the higher-biodiversity Valdivian temperate forests. Genus Nothofagus, also known as southern beech, is the predominant tree type and is characteristic of the biome. In fact, the presence of Nothofagus across the austral continents (southern Pacific Rim) is evidence that Antarctica, Australia, and South America were connected in geological time.

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This biome represents the southernmost forests in South America: It ranges through the southern Aysén and Magallanes regions of Chile; some western patches of Santa Cruz Province, Argentina; and to southern swaths of Tierra del Fuego, mainly from Lake Buenos Aires to Staten Island. The name refers to its proximity to the Strait of Magellan, which memorializes the global circumnavigation voyage of the Portuguese explorer Ferdinand Magellan in 1520.

Vegetation

The southern beech varieties found here include Nothofagus betuloides, N. antarctica, and N. pumilio. Generally, species richness is much lower than in the Valdivian forests to the north, but this unique biome supports relatively rich biodiversity in one of the harshest climates on the continent. Three plant community types are represented in the biome, mixed across ecotones and with neighboring biomes: moorlands, temperate forests, and deciduous forests. N. antarctica and N. pumilio are deciduous and typical of the eastern portion, while N. betuloides is evergreen and typical of the western zones.

At the highest elevations, this biome is dominated by permanent snow, ice caps, and glaciers. Generally speaking, it is characterized by cold temperatures, strong and permanent west winds, and high amounts of rain and snow. Landscapes are dramatic and often windswept, with few year-round human inhabitants.

The unique climate of the region has resulted in numerous endemic (found only here) plant species, but biodiversity is much lower than in surrounding biomes. Endemic plants include grasses such as Deschamsia kingii, Festucacirrosa, Poa darwiniana, and P. yaganica; and a suite of herbs and shrubs, including Onuris alismatifolia, Ourisia fuegiana, O.ruelloides, Senecio eightsii, S. humifusus, S. websteri, Nassauvia latissima, Acaena lucida, Perezia lactucoides, and Viola commersonii. Many plants are stunted because of the high winds; some are confined to leeward rock and other formations that provide shelter.

Fauna

Endemism in animals here is much lower than in plants. Much of the fauna occur marginally, and are more predominant in neighboring biomes. Among the birds are the Magellanic woodpecker (Campephilusmagellanicus), austral parakeet (Enicognathus ferrrugineus), and several near-endemic geese (Chloephaga hybrida and C. rubiceps).

Mammals have similar low diversity, but are notably visible in the landscape. Among the large mammals are puma (Puma concolor), two foxes (Pseudalopex griseus and P. culpaeus), guanaco (Lama guanicoe), Patagonian huemul (Hippocamelus bisulcus), southern pudu (Pudupudu), and southern river otter (Lontraprovocax). The only near-endemic mammals are grass mice such as Abrothrix (previously Akodon) hershkovitzi, A. lanosus, and A.markhami; currently, these are not considered to be threatened.

Environmental Threats

The Magellanic Subpolar Forests biome is seriously threatened by habitat conversion due to logging, human development in northern areas, and increases in toursim, grazing, and fires. In addition, the introduction of habitat-modifying exotic species such as the North American beaver (Castor canadensis) and the European rabbit (Oryctolagus cuniculus) have affected many plant communities. Beavers build dams and thus alter the habitat and soil characteristics, inhibiting forest restoration and damaging the original hydrological cycles.

Glaciers in the southern Patagonian region have been losing mass at a very rapid rate, one of the fastest glacier retreats recorded on Earth. Some of the permanent ice cover here has retreated up to 9 miles (14.5 kilometers) in the last 100 years—with some indications of recent acceleration. Besides more erratic spring runoff events, glacial shrinkage can lead to water shortages during drier months, affecting local habitat resilience and growth rates.

Climate change impacts upon this region have yet to be sharply predicted, but regional warming trends, precipitation disruption, or seasonal pattern shifts hold the potential to erode soils and challenge vegetation viability at different altitude gradients, and thus to fragment and otherwise stress the current habitat structure, food web, and fauna mix.

Bibliography

Brion, C. D., J. Puntieri Grigera, and E. Rapoport. “Plantas Exóticas en Bosques de Nothofagus. Comparaciones Preliminares Entre el Norte de la Patagonia y Tierra del Fuego.” Monograf Biologias de la Academia Nacional de Ciencias Exactas, Fisicas y Naturales 4, no. 1 (1988).

Davies, Bethan. "Shrinking Patagonian Glaciers." Antarctic Glaciers, 22 June 2020, www.antarcticglaciers.org/glaciers-and-climate/glacier-recession/shrinking-patagonian-glaciers/. Accessed 4 Dec. 2024.

De la Peña, M. R., and M. Rumboll. Birds of Southern South America and Antarctica. Princeton, NJ: Princeton University Press, 1998.

Gajardo, R. La Vegetación Natural de Chile, Clasificación y Distribución Geográfica. Santiago, Chile: Editorial Universitaria, 1994.

Lliboutry, Louis. “Glaciers of South America: Glaciers of Chile and Argentina.” In Williams, R. S. and J. G. Ferrigno, eds. Satellite Image Atlas of Glaciers of the World. Washington, DC: U.S. Geological Survey, 1998.

“The Magellanic Subpolar Forests: A Comprehensive Exploration of Nature’s Southern Frontier.” Latin American and Caribbean Geographic, 20 May 2024, lacgeo.com/magellanic-subpolar-forests. Accessed 4 Dec. 2024.

Redford, K. H., and J. F. Eisenberg. Mammals of the Neotropics 2, The Southern Cone. Chicago: University of Chicago Press, 1992.

Veblen, T. T., C. Donoso, T. Kitzberger, and A. J. Rebertus. “Ecology of Southern Chilean and Argentinean Nothofagus Forests.” In T. T. Veblen, R. S. Hill, and J. Read, eds., The Ecology and Biogeography of Nothofagus Forests. New Haven, CT: Yale University Press, 1996.

Villa-Martinez, Rodrigo, and Patricio I. Moreno. "Development and Resilience of Deciduous Nothofagus Forests Since the Last Glacial Termination and Deglaciation of the Central Patagonian Andes." Palaeogeography, Palaeoclimatology, Palaeoecology, vol. 574, 2021. DOI: 10.1016/j.palaeo.2021.110459. Accessed 4 Dec. 2024.