Dryosaurus

Kingdom: Animalia

Phylum: Chordata

Class: Reptilia

Order: Ornithischia

Family: Dryosauridae

Genus:Dryosaurus

Species:Dryosaurus altus

Introduction

Dryosaurus was a medium-sized herbivore that lived during the Late Jurassic period. Part of a large group of dinosaurs characterized by hoof-like feet and a bipedal stance, Dryosaurus was a relatively common herbivore that existed across a wide portion of the Northern Hemisphere.

In the Triassic period, the continents were united into a single landmass, known as Pangaea. During the Jurassic period, tectonic shifting and other geological forces caused the supercontinent to split into two smaller supercontinents, with Gondwana encompassing most of what would become the Southern Hemisphere landmass, and Laurasia, in the north, containing most of what would become North America, Europe, and parts of Asia.

Dryosaurus lived in the part of the supercontinent known as Western Laurasia. Fossils of a similar species have been found in Tanzania in East Africa, suggesting that an ancestral species migrated between Gondwana and Laurasia during the Jurassic period.

Classification

Biologists use two primary methods to classify organisms. Linnaean taxonomy groups organisms according to overall physical similarity and was used to create the basic categories of life used today, such as placing organisms into kingdoms, phyla, classes, etc. An alternate classification system, called cladistics, creates groups of animals that better reflect the evolutionary relationships between species. Cladistic analysis uses key characteristics to create “clades,” groups than contain organisms sharing a common characteristic derived from a common ancestor.

Using Linnaean taxonomy, paleontologists have placed Dryosaurus within the infraorder known as Ornithopoda, or “bird-footed” dinosaurs. These herbivorous dinosaurs had bony beaks and feet with three or four toes (in earlier forms). Dryosaurus, Valdosaurus, and Kangnasaurus species are relatively primitive members of the Ornithopoda group closely related to the hadrosaurs of the Cretaceous period.

Dryosaurus has been placed in the unranked clade iguanodontian, alongside dinosaurs such as Tenontosaurus. Members of this clade have specific teeth arrangements and are known for their enlarged nostrils.

117145960-101978.jpg

Anatomy

Examinations of Dryosaurus bones indicate that none of the specimens found have been fully grown, so paleontologists are uncertain of its maximum size. Specimens indicate a length of between 3 to 4 meters (9–13 ft) and a weight of around 90 kilograms (198 lbs).

Dryosaurus was believed to be largely bipedal but with strong, well-muscled arms. No skin impressions have been found for Dryosaurus, but it may have had pebbled or mosaic scale skin similar to its relatives, the hadrosaurs.

The skull of Dryosaurus was large with a short muzzle and a pronounced bill. Its cheeks contained rows of chewing teeth, and some paleontologists believe the cheek skin may have expanded to hold food. The animal had large eyes, relative to its skull size, indicating good vision.

Intelligence

Paleontologists have not calculated an EQ measurement for Dryosaurus, but other members of the Ornithopoda group generally fall in the range of 1.0 to 1.5, placing them on an intermediate level compared to other dinosaurs. Although some biologists consider EQ to be a poor measure of intelligence, many paleontologists believe that herbivores like Dryosaurus might not have been as intelligent as modern mammals occupying similar ecological roles.

117145960-101980.jpg

Reproduction and Population

Paleontologists have found Dryosaurus eggs and hatchlings at several fossil sites around the world. Hatchlings were about 22 centimeters (8 in) at birth, and bone analysis indicates that they could stand and walk shortly after hatching. Examinations of hatchling teeth indicate that the animals were capable of eating vegetation shortly after hatching.

Paleontologists are unaware if young Dryosaurus were left to their own after hatching, or whether parents remained with the young for protection. Discoveries of juvenile specimens in close association with larger specimens of several sizes may indicate that Dryosaurus utilized communal or herd breeding areas. As Dryosaurus is closely related to several species whose young required parental care, it is likely that Dryosaurus also displayed some degree of parental care, possibly remaining in family groups like some modern birds and mammals.

Diet

Paleontologists believe Dryosaurus was an herbivore, likely feeding on ferns and other low-growing vegetation. The animal had an enlarged abdominal cavity and may have had a large stomach in which vegetation could ferment, speeding vegetation.

Dryosaurus had a small, sharp beak used to crop leaves and stems from plants. Dryosaurus had rows of ridged teeth in its cheeks that would have been useful for grinding vegetation prior to swallowing. Paleontologists have theorized that Dryosaurus may have had expanded tissue in the cheek area, allowing it to hold partially masticated food before swallowing or to carry food while foraging.

Behavior

Based on its large eyes, Dryosaurus probably had good vision, which may have been an adaptation to help the animal avoid predators. Examinations of the legs also indicate that Dryosaurus could run at fast speeds, likely another adaptation to help avoid predators.

Fossil beds have revealed groups of Dryosaurus skeletons in close proximity, leading some paleontologists to believe that the animals travelled or congregated in groups. Alternately, groups of fossils may indicate chaotic gatherings, for example around a central food or water source.

Detailed examinations of young Dryosaurus indicate a larger head-to-body ratio that may have prevented it from walking on two legs, as adult Dryosaurus was believed to have done. Thus, young Dryosaurus was quadrupedal, walking on four legs until it reached a juvenile stage and assumed the bipedal stance of its parents.

Habitat and Other Life Forms

In the Late Jurassic, the Atlantic Ocean began to rise and separate Western Laurasia from Eastern Laurasia. The area that would become North America had diverse habitat, ranging from lush semi-tropical forests to dry, arid plains. Late Jurassic vegetation consisted primarily of gymnosperms, which include conifers and cycads, and members of the fern group.

There were a variety of small reptiles, amphibians, and mammals living in North America. As there were no birds or flying mammals, the air was filled with flying insects and flying reptiles called pterosaurs that were the dominant flying vertebrates throughout the Mesozoic era.

One of the most dominant herbivore groups in Jurassic North America was the sauropods, like Brachiosaurus, Diplodocus, and Apatosaurus. In addition, there were several species of herbivores like Stegosaurus that had long powerful tails, armed with spikes, and characteristic plates along their spines. Among the predators, there were large theropods, predatory dinosaurs like Torvosaurus and Ceratosaurus that probably hunted herbivores like Dryosaurus.

Research

Dryosaurus skeletons were first found in Dinosaur National Monument, a park spanning the borders of Colorado and Utah. The original species, Dryosaurusaltus, was named by paleontologist Othniel Marsh in 1894. Specimens have also been found in other parts of the western United States, indicating that the species was widespread. In 2018, several scientists proposed that specimens found in Dinosaur National Monument were actually the new species Dryosaurus elderae.

Fossils of a dinosaur sharing many physical similarities with Dryosaurus have been found in Tanzania. Paleontologists, however, noting minor skeletal differences, now refer to it as D. lettowvorbecki, which is in the same genus as Dryosaurus. Discovery of closely related species in North America and Africa indicate that the ancestor species spread when Gondwana and Laurasia were still connected by a passable land bridge.

117145960-101981.jpg

Bibliography

Briggs, Derek E. G., and Peter R. Crowther. Paleobiology II. Wiley-Blackwell, 2001.

Carpenter, Kenneth. Eggs, Nests and Baby Dinosaurs: A Look at Dinosaur Reproduction. Indiana UP, 1999.

Carpenter, Kenneth, and Peter M. Galton. "A Photo Documentation of Bipedal Ornithischian Dinosaurs from the Upper Jurassic Morrison Formation, USA." Geology of the Intermountain West, vol. 5, 2018, pp. 167–207, pdfs.semanticscholar.org/880a/79257a03d5d960e74221c3867ef9551b0fed.pdf. Accessed 5 May. 2020.

"Dinosaurs in their Time." Carnegie Museum of National History, carnegiemnh.org/explore/dinosaurs-in-their-time. Accessed 5 May. 2020.

"Dryosaurus." Natural History Museum, www.nhm.ac.uk/discover/dino-directory/dryosaurus.html. Accessed 5 May. 2020.

Fastovsky, David E., and David B. Weishampel. Dinosaurs: A Concise Natural History. Cambridge UP, 2009.

Fastovsky, David E., and David B. Evolution and Extinction of the Dinosaurs. Cambridge UP, 2005.

Horner, Jack, and James Gorman. How to Build a Dinosaur: Extinction Doesn’t Have to Be Forever. Dutton, 2009.

Horner, John R., and Edwin Dobb. Dinosaur Lives. Mariner Books, 1998.

Lucas, Spencer G. Dinosaurs: The Textbook. 4th ed., McGraw Hill, 2004.

Martin, Anthony J. Introduction to the Study of Dinosaurs. Blackwell, 2006.

"Paleoportal." UC Museum of Paleontology, University of California, Berkeley, ucmp.berkeley.edu . Accessed 2011.

Rage, Jean-Claude. “Gondwana, Tethys, and Terrestrial Vertebrates During the Mesozoic and Cenozoic.” Geological Society, London, Special Publications, vol. 37, Jan 1988, pp. 255–73.

Sellers, William Irvin, and Phillip Lars Manning. “Estimating Dinosaur Maximum Running Speeds Using Evolutionary Robotics.” Proceedings of the Royal Society of Biology, vol. 274, Nov 2007, pp. 2711–16.

Weishampel, David B., et al. Dinosauria. 3rd ed., University of California Press, 2007.